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Uncovering A Mass Extinction Event: The Lake Rototoa Mussel Survey

Water quality scientist Ebi Hussain led a Project Baseline citizen science team to determine what was happening to New Zealand’s Lake Rototoa mussel population. Crying cockles and mussels, alive, alive, No!! Here are the deets.

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By Ebrahim (Ebi) Hussain
Header photo courtesy of Oliver Horschig.

Lake Rototoa, a cold, monomictic1 dune lake in a rural area northwest of Auckland, New Zealand, is in peril. With a maximum depth of 26m/85 ft, Rototoa is the largest and deepest of a series of sand dune lakes along the country’s western coastline. Known for its increasingly rare, diverse population of native submerged macrophytes i.e., aquatic plants, and large, freshwater mussel beds, this lake is under increasing threat from a deteriorated water quality. Although the exact cause of this deterioration is unclear, the likely culprit is a combination of factors: eutrophication, land use activities, pest invasion, and climate change.

In late 2019, the Project Baseline Aotearoa Lakes team noted signs of a freshwater mussel population collapse as well as other evidence of environmental degradation. This was alarming, as freshwater mussels are rapidly declining in New Zealand, and globally, with 70 percent of the species considered at risk or threatened.

Many people are unaware that freshwater mussels are an important part of a lake ecosystem; as biofilters and bioturbators, they filter out nutrients, algae, bacteria, and fine organic material which helps purify the water. The loss of these keystone  species has likely contributed to the decline in water quality seen at Lake Rototoa.

The team’s observations prompted the design of a collaborative project between Project Baseline Aotearoa Lakes and the Auckland Council Biodiversity Team. This project is the first of its kind in New Zealand; it aims to fill critical knowledge gaps and, for the first time, quantify mussel populations in Lake Rototoa in a scientific manner. 

Ebrahim Hussain installing the transect. This peg marks the beginning of the first transect across the mussel bed. Photo by Oliver Horschig.

This project is the first of its kind in New Zealand; it aims to fill critical knowledge gaps and, for the first time, quantify mussel populations in Lake Rototoa in a scientific manner. 

The first objective was to assess the mussel population statistics, including species composition, abundance, size class, and recruitment success. The second objective was to determine habitat preferences, bed locations, and bed limiting factors. In order to satisfy the project objectives, the team designed a bespoke survey methodology to collect all the required information in a standardized way.

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Digging Into The Data

The initial series of dives focused on habitat mapping and collecting bed scale survey information. The team has mapped almost 5 km2/3.1 mi2 of lakebed and 2.2 km2/1.4 mi2 of mussel bed so far. This information provided critical insight into mussel bed formation and habitat preferences which the team used to inform the site selection for the more detailed follow up surveys. 

The first phase of surveys has been completed and the results are frightening. A total of 1604 mussels (Echyridella menziesii) were counted. The combined density across all three survey sites was 41.4 mussels per m2/3.8 mussels/ft2. Out of the 1604 mussels found, 1320 (82.3%) were dead and only 284 (17.7%) were alive. The dead mussel shells were in a similar condition to the live individuals indicating that they may have all died during a recent mass extinction event.

No juveniles were seen during the surveys and all the mussels were larger than 51 mm/2 in. The surveyed population is composed entirely of mature adults, 64.1% of live mussels were larger than 70 mm/2.8 in in length, 30.6% were between 61 to 70 mm/2.4 to 2.8 in and the remaining 5.3% were in the 51 to 60 mm/2 to 2.4 in size class.

Measuring an adult Echyridella menziesii, this individual is 78mm in length and approximately 15-20 years old. Photo by Oliver Horschig.

Individual dead mussels were not measured but were placed into approximate size classes, all dead mussels were larger than 51 mm/2 in with the majority of them being placed in the 61 to 70 mm and >70 mm size classes. The average age of the mussels surveyed was estimated to be between 20 and 30 years old based on their size. Some larger individuals were 80 to 100 mm long and were estimated to be around 50 years old.  

This aging population and lack of younger individuals indicates limited-to-no viable recruitment in the surveyed area for more than a decade. Considering that most of the live mussels were at the upper end of their life expectancy and that there was no evidence of recent recruitment, the long-term viability of the surveyed population is low. 

While the exact reasons for this population collapse are not known, recent lake surveys (fish, water quality, and macrophytes)  provide some indication of possible causes. Recent fish surveys indicate a significant drop in the number of  the primary intermediate host species. Both galaxiid and bully species are declining due to predation by pest fish species. Without these native fish, the mussels cannot effectively complete their life cycle.

The declining water quality of the lake is also a contributing factor. The lake’s change from an oligotrophic state, which is low in plant nutrients and high oxygen at depth, to a mesotrophic state with moderate nutrients, subjected it to increased eutrophication.  

Eutrophication causes an increase in bioavailable nutrients which stimulates algal growth and in turn causes high organic silting. This silt settles on the lakebed and decomposes creating areas of low dissolved oxygen, which can cause animal die offs. 

Some studies suggest that these mussels cannot survive at dissolved oxygen concentrations below 5mg/L and it is possible that the lake undergoes prolonged periods of low-dissolved oxygen during seasonal stratification. The wide scale coverage of benthic blue-green algal mats further points to periods of anoxia, or absence of oxygen, and general eutrophication. 



Mussel Mania

Due to the low nutrient concentrations and the filtration capacity of the extensive mussel population, Lake Rototoa historically had good water clarity. Mussel filtration rates generally match their food ingestion rate, but once they reach their food ingestion rate, no further filtration will occur. If there is a high concentration of food (phytoplankton and zooplankton) in the water, the filtration rate is likely to be low. This means that as the lake becomes more eutrophic, the algal biomass increases, and the mussel’s filtration rate will continue to decrease. 

An Echyridella menziesii bed, showing a mixture of live and dead adult sized mussels. Photo by Ebrahim Hussain.

This decrease in filtration rates will contribute to the declining visual clarity. The significant loss of mussel biomass and ultimately the loss of mussels in Lake Rototoa exacerbated the situation and may have facilitated a higher rate of eutrophication.

Sediment is also known to affect mussel populations, and there are signs of increased sedimentation; however, no clear evidence of smothering or suffocating was observed. The combination of the organic silt, sediment, and benthic algal growth can clog the mussel gills, so there are likely to be some sediment-induced population stressors.

In terms of bed extent and bed limiting factors, the team made several key observations. The mussels tended to prefer gentle slopes and did not occur in great densities on steep faced slopes/shelves. Water level, riparian vegetation extent, and wind/wave-induced disturbance appeared to dictate the upper extent. Mussel beds were generally established at a depth just below the permanent water line a short distance away from the end of the riparian edge. Fewer mussels were observed in shallow, exposed areas with visible signs of wind/wave-induced substrate disturbance. 

The establishment of aquatic plants, changes in substrate, thermoclines, and potentially anoxia limited the lower bed extent. Mussels were commonly found in lower numbers in amandaphyte stands within the wider bed area and were not found at all within dense charophyte meadows. Mussels tended to establish around isolated macrophyte stands rather than in them. The lower extent of the bed mirrored the start of the deeper charophyte meadows. The littoral zone had clearly defined sections of mussels in the shallower areas (1.5 to 5 m/5 to 16 ft ) and dense macrophyte dominated areas in the deeper portion (6 to 10 m), which were relatively devoid of mussels. 

Ebrahim Hussain counting and size classing mussels in a 0.25m 2 quadrate. Photo by Oliver Horschig.

In the absence of aquatic plants, the thermocline separating the warmer epilimnion above from the colder hypolimnion below appeared to dictate the lower bed extent. Almost no mussels were found past the thermocline, which was between 6 and 7 m/20 to 23 ft deep during the survey period. Since mussel bed establishment is not known to be thermally regulated, the limiting factor here may be anoxic conditions, commonly associated with hypolimnetic water. This assumption has not been validated, and a more detailed investigation of stratification profiles are planned for this upcoming year.

A clear limiting factor is the change in substrate seen past the 7 to 10 m/23 to 33 ft depth contour. The substrate changes from sand with a surficial layer of silt to a semi liquid silt/soft mud. No mussels or macrophytes were found in these areas, and the substrate does not appear to support bed establishment. Benthic algal mats covered the lower extent of some beds but did not clearly limit their establishment; since these mussels are mobile, presumably they will move if they are being smothered.

In Conclusion

Despite the concerning results, this project is a landmark event as it is the first study of its kind in New Zealand and the first detailed survey of the mussel population in Lake Rototoa. This project highlighted the pressures faced by our aquatic environments and exposed the ugly truth of what is going on below the surface. We have uncovered a mass extinction event that is currently occurring in our back yard that no one even knew was happening. 

We have uncovered a mass extinction event that is currently occurring in our back yard that no one even knew was happening.

Now more than ever, projects like this are critical. Our environments are under increasing pressure, and it is up to all of us to take action to ensure that we preserve these ecosystems for future generations.

The follow-up phases of this project are planned to be carried out this summer. The data we have collected thus far has enabled the Auckland Council to make informed decisions on how best to manage these threatened species and preserve native biodiversity. We hope that our continued efforts at this lake will contribute to preserving this ecosystem and prevent the complete extinction of these threatened species.

Footnotes:

  1. Cold monomictic lakes are lakes that are covered by ice throughout much of the year. During their brief “summer”, the surface waters remain at or below 4°C. The ice prevents these lakes from mixing in winter. During summer, these lakes lack significant thermal stratification, and they mix thoroughly from top to bottom. These lakes are typical of cold-climate regions. 

Dive Deeper:

InDepth V 1.6: Bringing Citizen Science To Lake Pupuke by Ebrahim Hussain

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Ebrahim (Ebi) Hussain is a water quality scientist who grew up in South Africa. As far back as he can remember he has always wanted to scuba dive and explore the underwater world. He began diving when he was 12 years old and he has never looked back. Diving opened up a new world for him and he quickly developed a passion for aquatic ecosystems and how they work. The complexity of all the abiotic and biotic interactions fascinates him and has inspired Ebi to pursue a career in this field.
He studied aquatic ecotoxicology and zoology at university, and it was clear that Ebi wanted to spend his life studying these subsurface ecosystems and the anthropogenic stressors that impact them. After traveling to New Zealand, Ebi decided to move to this amazing country. The natural beauty drew him in, and even though there were signs of environmental degradation, there was still hope. Ebi founded Project Baseline Aotearoa Lakes with the goal of contributing to preserving and enhancing this natural beauty as well as encouraging others to get involved in actively monitoring their natural surroundings.

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Twenty-five Years in the Pursuit of Excellence – The Evolution and Future of GUE

Founder and president Jarrod Jablonski describes his more than a quarter of a century long quest to promote excellence in technical diving.

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by Jarrod Jablonski. Images courtesy of J. Jablonski and GUE unless noted.

The most difficult challenges we confront in our lives are the most formative and are instrumental in shaping the person we become. When I founded Global Underwater Explorers (GUE), the younger version of myself could not have foreseen all the challenges I would face, but equally true is that he would not have known the joy, the cherished relationships, the sense of purpose, the rich adventures, the humbling expressions of appreciation from those impacted, or the satisfaction of seeing the organization evolve and reshape our industry. Many kindred souls and extraordinary events have shaped these last 25 years, and an annotated chronology of GUE is included in this issue of InDEPTH. This timeline, however, will fail to capture the heart behind the creation of GUE, it will miss the passionate determination currently directing GUE, or the committed dedication ready to guide the next 25 years.

Photo courtesy of Kirill Egorov

I don’t remember a time that I was not in, around, and under the water. Having learned to swim before I could walk, my mother helped infuse a deep connection to the aquatic world. I was scuba certified in South Florida with my father, and promptly took all our gear to North Florida where I became a dive instructor at the University of Florida. It was then that I began my infatuation with cave diving. I was in the perfect place for it, and my insatiable curiosity was multiplied while exploring new environments. I found myself with a strong desire to visit unique and hard-to-reach places, be they far inside a cave or deep within the ocean. 

My enthusiasm for learning was pressed into service as an educator, and I became enamored with sharing these special environments. Along with this desire to share the beauty and uniqueness of underwater caves was a focused wish to assist people in acquiring the skills I could see they needed to support their personal diving goals. It could be said that these early experiences were the seeds that would germinate, grow, mature, and bloom into the organizing principles for GUE.

Brent Scarabin, Jarrod and George “Trey” Irvine getting ready to dive.
Jarrod with his Halcyon PVR-BASC prototype.
George Irvine and Jarrod conducting the original DIR workshop.

The Pre-GUE Years

Before jumping into the formational days of GUE, allow me to help you visualize the environment that was the incubator for the idea that became GUE’s reality. By the mid-1990s, I was deeply involved in a variety of exploration activities and had been striving to refine my own teaching capacity alongside this growing obsession for exploratory diving. While teaching my open water students, I was in the habit of practicing to refine my own trim and buoyancy, noticing that the students quickly progressed and were mostly able to copy my position in the water. Rather than jump immediately into the skills that were prescribed, I started to take more time to refine their comfort and general competency. This subtle shift made a world of difference in the training outcomes, creating impressive divers with only slightly more time and a shift in focus. In fact, the local dive boats would often stare in disbelief when told these divers were freshly certified, saying they looked better than most open water instructors! 

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By this point in my career, I could see the problems I was confronting were more systemic and less individualistic. In retrospect, it seemed obvious that key principles had been missing in both my recreational and technical education, not to mention the instructor training I received. The lack of basic skill refinement seemed to occur at all levels of training, from the beginner to the advanced diver. Core skills like buoyancy or in-water control were mainly left for divers to figure out on their own and almost nobody had a meaningful emphasis on efficient movement in the water. It was nearly unheard of to fail people in scuba diving, and even delaying certification for people with weak skills was very unusual. This remains all too common to this day, but I believe GUE has shifted the focus in important ways, encouraging people to think of certification more as a process and less as a right granted to them because they paid for training. 

L2R: Jarrod Todd Kincaid and Rickard Lundgren plotting their 1999 Britannic expedition.

The weakness in skill refinement during dive training was further amplified by little-to-no training in how to handle problems when they developed while diving, as they always do. In those days, even technical/cave training had very little in the way of realistic training in problem resolution. The rare practice of failures was deeply disconnected from reality. For example, there was almost no realistic scenario training for things like a failed regulator or light. What little practice there was wasn’t integrated into the actual dive and seemed largely useless in preparing for real problems. I began testing some of my students with mock equipment failures, and I was shocked at how poorly even the best students performed. They were able to quickly develop the needed skills, but seeing how badly most handled their first attempts left me troubled about the response of most certified divers should they experience problems while diving, as they inevitably would. 

Diving Fatalities

Meanwhile, I was surrounded by a continual progression of diving fatalities, and most appeared entirely preventable. The loss of dear friends and close associates had a deep impact on my view of dive training and especially on the procedures being emphasized at that time within the community. The industry, in those early days, was wholly focused on deep air and solo diving. However, alarmingly lacking were clear bottle marking or gas switching protocols. It seemed to me to be no coincidence that diver after diver lost their lives simply because they breathed the wrong bottle at depth. Many others died mysteriously during solo dives or while deep diving with air. 

One of the more impactful fatalities was Bob McGuire, who was a drill sergeant, friend, and occasional dive buddy. He was normally very careful and focused. One day a small problem with one regulator caused him to switch regulators before getting in the water. He was using a system that used color-coded regulators to identify the gas breathed. When switching the broken regulator, he either did not remember or did not have an appropriately colored regulator. This small mistake cost him his life. I clearly remember turning that one around in my head quite a bit. Something that trivial should not result in the loss of a life. 

Also disturbing was the double fatality of good friends, Chris and Chrissy Rouse, who lost their lives while diving a German U-boat in 70 m/230 ft of water off the coast of New Jersey. I remember, as if the conversation with Chris were yesterday, asking him not to use air and even offering to support the cost as a counter to his argument about the cost of helium. And the tragedies continued: The loss of one of my closest friends Sherwood Schille, the death of my friend Steve Berman who lived next to me and with whom I had dived hundreds of times, the shock of losing pioneering explorer Sheck Exley, the regular stream of tech divers, and the half dozen body recoveries I made over only a couple years, which not only saddened me greatly, but also made me angry. Clearly, a radically different approach was needed.

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Learning to Explore

Meanwhile, my own exploration activities were expanding rapidly. Our teams were seeking every opportunity to grow their capability while reducing unnecessary risk. To that end, we ceased deep air diving and instituted a series of common protocols with standardized equipment configurations, both of which showed great promise in expanding safety, efficiency, and comfort. We got a lot of things wrong and experienced enough near misses to keep us sharp and in search of continual improvement. 

Casey McKinlay and Jarrod with stages and Gavin scooters in Wakulla Springs. Photo courtesy of David Rhea

But we looked carefully at every aspect of our diving, seeking ways to advance safety, efficiency, and all-around competency while focusing plenty of attention into the uncommon practice of large-scale, team diving, utilizing setup dives, safety divers, and inwater support. We developed diver propulsion vehicle (DPV) towing techniques, which is something that had not been done previously. We mostly ignored and then rewrote CNS oxygen toxicity calculations, developed novel strategies for calculating decompression time, and created and refined standard procedures for everything from bottle switching to equipment configurations. Many of these developments arose from simple necessity. There were no available decompression programs and no decompression tables available for the dives we were doing. Commonly used calculations designed to reduce the risk of oxygen toxicity were useless to our teams, because even our more casual dives were 10, 20, or even 30 times the allowable limit. The industry today takes most of this for granted, but in the early days of technical diving, we had very few tools, save a deep motivation to go where no one had gone before.

All in a dive of diving for the WKPP.

Many of these adventures included friends in the Woodville Karst Plain Project (WKPP), where I refined policies within the team and most directly with longtime dive buddy George Irvine. This “Doing it Right” (DIR) approach sought to create a more expansive system than Hogarthian diving, which itself had been born in the early years of the WKPP and was named after William Hogarth Main, a friend and frequent dive buddy of the time. By this point, I had been writing about and expanding upon Hogarthian diving for many years. More and more of the ideas we wanted to develop were not Bill Main’s priorities and lumping them into his namesake became impractical, especially given all the debate within the community over what was and was not Hogarthian. 

A similar move from DIR occurred some years later when GUE stepped away from the circular debates that sought to explain DIR and embraced a GUE configuration with standard protocols, something entirely within our scope to define.

These accumulating events reached critical mass in 1998. I had experienced strong resistance to any form of standardization, even having been asked to join a special meeting of the board of directors (BOD) for a prominent cave diving agency. Their intention was to discourage me from using any form of standard configuration, claiming that students should be allowed to do whatever they “felt’ was best. It was disconcerting for me, as a young instructor, to be challenged by pioneers in the sport; nevertheless, I couldn’t agree with the edict that someone who was doing something for the first time should be tasked with determining how it should be done. 

This sort of discussion was common, but the final straw occurred when I was approached by the head of a technical diving agency, an organization for which I had taught for many years. I was informed that he considered it a violation of standards not to teach air to a depth of at least 57 m/190 ft. This same individual told me that I had to stop using MOD bottle markings and fall in line with the other practices endorsed by his agency. Push had finally come to shove, and I set out to legitimize the training methods and dive protocols that had been incubating in my mind and refined with our teams over the previous decade. Years of trial and many errors while operating in dynamic and challenging environments were helping us to identify what practices were most successful in support of excellence, safety, and enjoyment.

Forming GUE

Forming GUE as a non-profit company was intended to neutralize the profit motivations that appeared to plague other agencies. We hoped to remove the incentive to train—and certify—the greatest number of divers as quickly as possible because it seemed at odds with ensuring comfortable and capable divers. The absence of a profit motive complemented the aspirational plans that longtime friend Todd Kincaid and I had dreamed of. We imagined a global organization that would facilitate the efforts of underwater explorers while supporting scientific research and conservation initiatives. 

I hoped to create an agency that placed most of the revenue in the hands of fully engaged and enthusiastic instructors, allowing them the chance to earn a good living and become professionals who might stay within the industry over many years. Of course, that required forgoing the personal benefit of ownership and reduced the revenue available to the agency, braking its growth and complicating expansion plans. This not only slowed growth but provided huge challenges in developing a proper support network while creating the agency I envisioned. There were years of stressful days and nights because of the need to forgo compensation and the deep dependance upon generous volunteers who had to fit GUE into their busy lives. If it were not for these individuals and our loyal members, we would likely never have been successful. Volunteer support and GUE membership have been and remain critical to the growing success of our agency. If you are now or have ever been a volunteer or GUE member, your contribution is a significant part of our success, and we thank you. 

Photo courtesy of Kirill Egorov

The challenges of the early years gave way to steady progress—always slower than desired, with ups and downs, but progress, nonetheless. Some challenges were not obvious at the outset. For example, many regions around the world were very poorly developed in technical diving. Agencies intent on growth seemed to ignore that problem, choosing whoever was available, and regardless of their experience in the discipline, they would soon be teaching. 

This decision to promote people with limited experience became especially problematic when it came to Instructor Trainers. People with almost no experience in something like trimix diving were qualifying trimix instructors. Watching this play out in agency after agency, and on continent after continent, was a troubling affair. Conversely, it took many years for GUE to develop and train people of appropriate experience, especially when looking to critical roles, including high-level tech and instructor trainers. At the same time, GUE’s efforts shaped the industry in no small fashion as agencies began to model their programs after GUE’s training protocols. Initially, having insisted that nobody would take something like Fundamentals, every agency followed suit in developing their own version of these programs, usually taught by divers that had followed GUE training. 

This evolving trend wasn’t without complexity but was largely a positive outcome. Agencies soon focused on fundamental skills, incorporated some form of problem-resolution training, adhered to GUE bottle and gas switching protocols, reduced insistence on deep air, and started talking more about developing skilled divers, among other changes. This evolution was significant when compared to the days of arguing about why a person could not learn to use trimix until they were good while diving deep on air. 

To be sure, a good share of these changes was more about maintaining business relevance than making substantive improvements. The changes themselves were often more style than substance, lacking objective performance standards and the appropriate retraining of instructors. Despite these weaknesses, they remain positive developments. Talking about something is an important first step and, in all cases, it makes room for strong instructors in any given agency to practice what is being preached. In fact, these evolving trends have allowed GUE to now push further in the effort to create skilled and experienced divers, enhancing our ability to run progressively more elaborate projects with increasingly more sophisticated outcomes. 

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The Future of GUE

The coming decades of GUE’s future appear very bright. Slow but steady growth has now placed the organization in a position to make wise investments, ensuring a vibrant and integrated approach. Meanwhile, evolving technology and a broad global base place GUE in a unique and formidable position. Key structural and personnel adjustments complement a growing range of virtual tools, enabling our diverse communities and representatives to collaborate and advance projects in a way that, prior to now, was not possible. Strong local communities can be easily connected with coordinated global missions; these activities include ever-more- sophisticated underwater initiatives as well as structural changes within the GUE ecosystem. One such forward-thinking project leverages AI-enabled, adaptive learning platforms to enhance both the quality and efficiency of GUE education. Most agencies, including GUE, have been using some form of online training for years, but GUE is taking big steps to reinvent the quality and efficiency of this form of training. This is not to replace, but rather to extend and augment inwater and in-person learning outcomes. Related tools further improve the fluidity, allowing GUE to seamlessly connect previously distant communities, enabling technology, training, and passion to notably expand our ability to realize our broad, global mission.

Photo courtesy of Kirill Egorov

Meanwhile, GUE and its range of global communities are utilizing evolving technologies to significantly expand the quality and scope of their project initiatives. Comparing the impressive capability of current GUE communities with those of our early years shows a radical and important shift, allowing results equal or even well beyond those possible when compared even with well-funded commercial projects. Coupled with GUE training and procedural support, these ongoing augmentations place our communities at the forefront of underwater research and conservation. This situation will only expand and be further enriched with the use of evolving technology and closely linked communities. Recent and planned expansions to our training programs present a host of important tools that will continue being refined in the years to come. Efforts to expand and improve upon the support provided to GUE projects with technology, people, and resources are now coming online and will undoubtedly be an important part of our evolving future.

The coming decades will undoubtedly present challenges. But I have no doubt that together we will not only overcome those obstacles but we will continue to thrive. I believe that GUE’s trajectory remains overwhelmingly positive, for we are an organization that is continually evolving—driven by a spirit of adventure, encouraged by your heartwarming stories, and inspired by the satisfaction of overcoming complex problems. Twenty-five years ago, when I took the path less traveled, the vision I had for GUE was admittedly ambitious. The reality, however, has exceeded anything I could have imagined. I know that GUE will never reach a point when it is complete but that it will be an exciting lifelong journey, one that, for me, will define a life well lived. I look forward our mutual ongoing “Quest for Excellence.”

See Listings Below For Additional Resources On GUE And GUE Diving!

Jarrod is an avid explorer, researcher, author, and instructor who teaches and dives in oceans and caves around the world. Trained as a geologist, Jarrod is the founder and president of GUE and CEO of Halcyon and Extreme Exposure while remaining active in conservation, exploration, and filming projects worldwide. His explorations regularly place him in the most remote locations in the world, including numerous world record cave dives with total immersions near 30 hours. Jarrod is also an author with dozens of publications, including three books.

A Few GUE Fundamentals

Similar to military, commercial and public safety divers, Global Underwater Explorers (GUE) is a standards-based diving community, with specific protocols, standard operating procedures (SOPs) and tools. Here are selected InDEPTH stories on some of the key aspects of GUE diving, including a four-part series on the history and development of GUE decompression procedures by founder and president Jarod Jablonski.

Anatomy of a Fundamentals Class

GUE Instructor Examiner Guy Shockey explains the thought and details that goes into GUE’s most popular course, Fundamentals, aka “Fundies,” which has been taken by numerous industry luminaries. Why all the fanfare? Shockey characterizes the magic as “simple things done precisely!

Back to Fundamentals: An Introduction to GUE’s Most Popular Diving Course

Instructor evaluator Rich Walker attempts to answer the question, “why is Fundamentals GUE’s most popular diving course?” Along the way, he clarifies some of the myths and misconceptions about GUE training. Hint: there is no Kool-Aid. 

The GUE Pre-dive Sequence

As you’d expect, Global Underwater Explorers (GUE) has a standardized approach to prepare your equipment for the dive, and its own pre-dive checklist: the GUE EDGE. Here explorer and filmmaker Dimitris Fifis preps you to take the plunge, GUE-style.

The Flexibility of Standard Operating Procedures

Instructor trainer Guy Shockey discusses the purpose, value, and yes, flexibility of standard operating procedures, or SOPs, in diving. Sound like an oxymoron? Shockey explains how SOPs can help offload some of our internal processing and situational awareness, so we can focus on the important part of the dive—having FUN!

Standard Gases: The Simplicity of Everyone Singing the Same Song

Like the military and commercial diving communities before them, Global Underwater Explorers (GUE) uses standardized breathing mixtures for various depth ranges and for decompression. Here British wrecker and instructor evaluator Rich Walker gets lyrical and presents the reasoning behind standard mixes and their advantages, compared with a “best mix” approach. Don’t worry, you won’t need your hymnal, though Walker may have you singing some blues.

Rules of Thumb: The Mysteries of Ratio Deco Revealed

Is it a secret algorithm developed by the WKPP to get you out of the water faster sans DCI, or an unsubstantiated decompression speculation promoted by Kool-Aid swilling quacks and charlatans? British tech instructor/instructor evaluator Rich Walker divulges the arcane mysteries behind GUE’s ratio decompression protocols in this first of a two part series.

The Thought Process Behind GUE’s CCR Configuration

Global Underwater Explorers is known for taking its own holistic approach to gear configuration. Here GUE board member and Instructor Trainer Richard Lundgren explains the reasoning behind its unique closed-circuit rebreather configuration. It’s all about the gas!

GUE and the Future of Open Circuit Tech Diving

Though they were late to the party, Global Underwater Explorers (GUE) is leaning forward on rebreathers, and members are following suit. So what’s to become of their open circuit-based TECH 2 course? InDepth’s Ashley Stewart has the deets.

Project Divers Are We

Diving projects, or expeditions—think Bill Stone’s Wakulla Springs 1987 project, or the original explorations of the Woodville Karst Plain’s Project (WKPP)—helped give birth to technical diving, and today continue as an important focal point and organizing principle for communities like Global Underwater Explorers (GUE). The organization this year unveiled a new Project Diver program, intended to elevate “community-led project dives to an entirely new level of sophistication.” Here, authors Guy Shockey and Francesco Cameli discuss the power of projects and take us behind the scenes of the new program

Decompression, Deep Stops and the Pursuit of Precision in a Complex World In this first of a four-part series, Global Underwater Explorers’ (GUE) founder and president Jarrod Jablonski explores the historical development of GUE decompression protocols, with a focus on technical diving and the evolving trends in decompression research.

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